On the first reliable find of the genus Tholodus (Reptilia: Ichthyopterygia) in the Asian peripheral area of the Panthalassic ocean


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Until recently, Tholodus, a rare ichthyosaur with spherical tooth crowns of the pressing–crushing type, has only been known from the Middle Triassic (Anisian) beds of Western Europe. A record of Tholodus remains in the Karazin Formation (Middle Anisian) of southern Primorye (Russian Far East) significantly expands the range of these reptiles.

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M. Arkhangelsky

Ecological Faculty; Geological Faculty

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N. Zverkov

Geological Faculty

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Yu. Zakharov

Far East Geological Institute, Far East Division

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I. Borisov

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