


No 3 (2025)
RESEARCH
The concepts of sustainability and climax of native spruce forests from the standpoint of successional dynamics of the forest community
Abstract
The concepts of climax and sustainability of forest communities were considered using forests of native spruce formations of the taiga zone of European Russia as an example. Objects and methods. Native spruce forests of different ages, different dynamic states and evolutionary formations were selected by a random sample from the general database of spruce communities of the taiga zone of the European North. A total of 30 spruce forests from all taiga subzones were selected. Results and discussion. The structure of age series and analysis of different generations’ trees volumes in forest stands were studied to determine the successional position of spruce forests. Biogeocenoses with no more than two excesses of tree volumes (50 % or more of the forest stand wood stock) in adjacent age generations cannot be considered forests of the climax dynamic state, but are treated as communities of transitional phases of dynamics. Spruce forests with relatively uniform distribution of tree volumes within generations of adjacent positions in the age series of the forest stand correspond to a stable state to the greatest extent, which significantly increases the time period of their stable structural position, close to the climax state. Statistical analysis of the relationship between age and volume of trees in biogeocenoses of different successional positions confirms these expectations: the relationship between predictors is “high, close” — R2 > 0.8. Conclusion. A forest community can reach a climax stable state with a relatively uniform distribution of tree volumes in age generations, constituting more than half of the age series of the forest stand.



Crown’s and Canopy’s Morphometry-Based Estimation Models for the Tree’sand Stand’s Trunk Diameter of Forest-Forming Species of Northern Eurasia Available for Lidar Scanning
Abstract
Within the framework of climate-smart forestry, accurate information on phytomass and carbon deposition capacity of forests is necessary. To date, many empirical models have been published and many taxation standards have been compiled to assess the trees’ and stands’ phytomass based on morphometric indicators measured in sample plots. However, using them in order to assess the carbon-depositing capacity of forests across large areas by means of traditional land-based forest inventory is quite a laborious undertaking. The laser (lidar) technology may be used as an alternative, but it does not allow determining the main mass-forming parameters — the tree trunk’s diameters or the average trunk diameter within the stand. To combine traditional empirical models and tables of phytomass with remote sensing data, intermediate models are needed to estimate the diameter of the tree trunk or the average trunk diameter within the stand, depending on the morphometry of the canopy, recorded either by terrestrial methods or remotely. The purpose of this study was to design the models of the tree trunk’s diameter’s dependence and the average trunk diameter within the stand on the main morphometric indicators of the canopy, obtained by ground measurements, but also available for lidar scanning. The models are constructed at the level of genera as aggregates of vicarious species. The materials of two previously compiled databases on phytomass and morphometric structure of 5320 trees and 5817 stands of Eurasia were used as the initial data for the study. Two-factor allometric dependences were constructed for 13 genera: (a) the stem diameter at breast height relation to the height of the tree and the diameter of the crown and (b) the average trunk diameter of the stand relation to the average tree height and stand’s density, explaining in most cases from 90 to 97% of the dependent variable’s variability. The proposed models based on traditional ground-based taxation data can be directly applied in lidar technologies or used to validate models based on lidar sensing data. This is especially important due to the lack of ground-based measurements of tree and stand morphometry for most existing species and habitats. The use of the proposed models based on the results of remote registration of crown and canopy morphology makes it possible to assess the phytomass and carbon pool of trees and stands in some territories in real time by combining them with available standards and specifications for determining the phytomass of trees and stands.



Definition Criteria and Structure of Stands in the Old-Growth Forests of the Altai Mountainsy
Abstract
Old-growth forests with very old trees are important ecosystems for nature and humans. There are practically no publications regarding their structure in the Altai Mountains. The objective of the study is to identify old-growth forests in the Altai Mountains using structural and successional criteria of the old-growth stage based on the analysis of the structure of their tree storey. In these forests, 29 sample plots were laid in 8 study areas on 4 mountain ranges. Their tree storey is dominated by Siberian pine and Siberian larch. The average maximum age of pine was 410—590 years, larch — 360—560 years, the average age of the main generation of trees was equal or less. These uneven-aged stands are morphologically complex, with 1—3 tree storeys and 2—4 generations of trees. Ontogenetic spectra of the Siberian pine coenopopulation are complete, while those of the larch are incomplete. Siberian pine dominates as seedlings. Age distributions of tree coenopopulations have an inverse J-shaped, uneven, multimodal form. A distinctive feature of these forests is the presence among the age parcels (gap mosaics) the parcels with dominance of the g3 (g3+g3–ss+g3–s) state individuals, with the possibility of their further transition to individuals of other growth forms: “livingstump” or “living log”. Due to the latter, it is possible to extend the ontogenesis of individual trees by another ≈200 years. And the complete ontogenesis of a tree with further transition to another growth form can be 700—800 years or more. In the absence of catastrophic impacts, old-growth forests of Siberian pine and larch can exist indefinitely. Criteria for old-growth forests in the Altai Mountains have been clarified.



Evapotranspiration in northern Taiga forests of the Nizhnyaya Tunguska river basin factoring in their post-fire dynamics
Abstract
The paper highlights the impact of forest cover disturbance on evapotranspiration changes within a river basin., The dynamics of forest cover disturbance as a result of forest fires in the Nizhnyaya Tunguska River basin for 2001—2020 was analysed based on Landsat satellite imagery. It was found that, as regards the spatial aspect, evapotranspiration in the Nizhnyaya Tunguska River basin is differentiated between the test catchments, which differ in climatic conditions and vegetation cover. Concerning the temporal aspect, a decrease in the weighted average evapotranspiration was noted as the area of burnt areas accumulated. It was revealed that the difference in the evapotranspiration decrease in undisturbed and disturbed areas depends both on the overall disturbance of the forest cover in the catchment area and on the area of burnt areas from the current year, with the effect of fresh burnt areas showing if their area is more than 3% of the catchment area.



Radial growth of Dahurian larch at the upper altitudinal limit in the western part of the Putorana Plateau
Abstract
Studying the growth of woody plants is extremely important due to the need to assess the response of vegetation to the ongoing climate change, especially in the mountainous regions of the Subarctic, where an increase in surface air temperature has been observed in recent decades. This paper presents the results of a study of the radial growth patterns of Dahurian larch (Larix gmelinii (Rupr.) Rupr.) at four altitudinal levels of the forest – mountain tundra ecotone on the Putorana Plateau slopes of western and eastern exposures. A comparative analysis of the radial growth magnitude and dynamics of trees belonging to 4 different age groups in forest stands of different density and altitudinal position was carried out. It was found that the radial increment’s sensitivity was higher in older trees, indicating that climatic factors had the greatest impact on their growth. It was discovered that young trees of the same age at higher hypsometric levels had higher morphometric characteristics values compared to those growing down the slope in more closed-up stands. The results of the study showed that intraspecific competition under these conditions had a greater impact on the growth of young trees than climatic factors. It was found that, while the climatic conditions in the study area underwent general improvement, no increase in the width of annual rings was observed in adult trees in recent decades. It was found that morphometric parameters and the width of annual rings in young trees had greater values on slopes of the eastern rather than the western exposure.



Polyvariance of the Siberian fir’s undergrowth development in dark coniferous forests of the Northern user
Abstract
Knowing the patterns of undergrowth development in forest-forming tree species is necessary for the correct reforestation assessment. The objective of the study was to characterise the of Siberian fir undergrowth development using the biological age concept and to assess the undergrowth’s prospects at different vitality. Based on the analysis of morphometric indices and the linear growth’s course of model Siberian fir plants (up to 3.5 m in height), diagnostic features of the ontogenetic stages im1, im2, v1 and vitality levels in the last two stages were formulated. To assess the model plants’ prospects, their height increment was predicted using the Mitscherlich function and the prediction results were compared to the height and age of generative trees grown in the same conditions. It was shown that to achieve the height of generative trees at a comparable age, the undergrowth should have an average height increment of at least 15—20 cm/year — typical values for plants of normal vitality on im2 and v1 ontogenetic stages; these plants are classified as high-prospects undergrowth. Another most clear diagnostic feature of the high-prospects undergrowth’s external structure is the presence of at least 2 living branches on all annual trunk increments. Undergrowth of the im2 (normal and low vitality) and v1 (low vitality) ontogenetic stages is classified as medium-prospects, and low vitality undergrowth of im2 and v1 ontogenetic stages is classified as questionable with regards to prospects. The undergrowth on the ontogenetic stage im1 hasn’t developed sufficiently to assess its prospects and the differentiation by vitality is not yet clearly expressed.



Efficiency of prolonged-release mineral fertilisers when growing Scots pine seedlings with closed root systems in cell packs
Abstract
In Russia, given the observed trend of increasing demand for growing high-quality planting material of year-old Scots pine (Pinus sylvestris L.) seedlings with a closed root system, an increase in the planned share of them planted during reforestation up to at least 30% of the total volume comes off as natural. The objective of the study is to analyse starter mineral treatment via the means of prolonged-release fertilisers and to assess their effectiveness for growing year-old seedlings of P. sylvestris as regards their development rate and changes in their inventory and biometric indicators. The methods used were agrochemical analysis of soil, manual biometric counting of seedlings and statistical analysis. The object of the study is starter mineral treatment introduced into the cell packs with wet high-moor peat of high acidity. Starter mineral treatment using urea capsules allows achieving 98% seed germination, reducing the rate of calendar development of year-old P. sylvestris seedlings from the seeds of the XI forest seed region until they reach standard parameters with less thinning. Growing P. sylvestris seedlings with the introduction of potassium monophosphate, phosphate rock, encapsulated urea in doses of 0,0369, 3,7647, 0,1159 kg/m3, respectively, allows seedlings to reach the largest root collar diameter of up to 2,2±0,4 mm before the age of one year. The use of encapsulated urea with hydroquinone for a total dose of 0,1162 kg/m3 stimulates the growth of seedlings in stem height the most, reaching 17,9±1,4 cm. In the experimental variants, unlike in the control with “Aquarin 5” fertiliser and a 0,5 cm layer of pine sawdust, the formation of verticils was noted. The experimental variant using urea capsules is the most effective variant of starter application of mineral treatments. It allows to reach a stem height of 16,8±1,4 cm and a root collar diameter of 2,2±0,4 mm with the lowest thinning percentage equal to 16,7% of the control, up to 38,3%, after their random sampling for agrochemical analysis and removal of dead seedlings.



Optimal and critical climatic seed transfer distance of Scots pine in the coniferous-broadleaf forests zone
Abstract
To maintain forest productivity and conservation, it is necessary to ensure the transfer of seeds for the creation of forest plantations in accordance with the scale and direction of climate change. The aim of this study was to analyse historical data on the growth and survival of Scots pine (Pinus sylvestris L.) of various provenances to assess the optimal and critical climatic seed transfer distance to the coniferous-broadleaf forests of the European Russia. The study was based on data on average heights and diameters at 17 years and tree survival at 5 years for 54 pine populations from the East European Plain and south-eastern Europe in plantations of the Kovrovsky district of the Vladimir region. The effect of genotype on phenotypic parameters of populations was estimated using a transfer function that included ecological distances of the number of days with average daily temperatures exceeding 5°C and annual precipitation. Under past and current climate conditions, seeds of local populations can be recommended for planting. By 2041—2070, according to model estimates, there will be a shift in the optimal areas of seed transfer in the south-western direction, thus, as a result of rapid climate change, local populations will show a decrease in productivity relative to those transferred from other areas and having better adaptation to new conditions. Dynamic forest seed zoning has been proposed as one of the tools for implementing the principles of climate-smart forestry in Russia.



OBITUARY
ПАМЯТИ ОЛЬГИ ВСЕВОЛОДОВНЫ СМИРНОВОЙ (09.10.1939–30.12.2024)


